We don't want a "species" sense of range, and a "behavioral" sense and an "epidemiological" sense.
the sort of axiom we need for this is as follows: (20) (∀ x, y, z)range (x, y, z)  x ranges from y to z if and only if there is a scale s with a subscale s 1 whose bottom is y and whose top is z, such that some member u 1 of x is at y, some member u 2 of x is at z, and every member u of x is at some point v in s 1 .